The study looked at how different environments affect the levels of iron and zinc in wheat grains, as well as the relationship between these micronutrients and grain yield. They found that wheat grains are naturally low in iron and zinc, making them important targets for biofortification. The researchers discovered that grain iron and zinc concentrations are positively correlated, but their relationship with yield components varies depending on the environment. This means that breeding for higher iron and zinc levels in wheat grains can be done without negatively impacting grain yield.